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Which Constellation carried a rotating radome like the E-3 Sentry's?
Short answer
The WV-2E, an experimental version of the US Navy's WV-2 Warning Star modified to carry a rotating radar dome similar to the Boeing E-3 Sentry's. It was later redesignated EC-121L, anticipating by decades the typical silhouette of airborne early warning aircraft.

The starting Warning Star
The United States Navy commissioned Lockheed to build a long-range flying radar based on the L-1049 Super Constellation. It was first designated PO-2W and then WV-2 Warning Star, powered by Wright R-3350-34 or R-3350-42 engines rated at 3,400 hp (2,536 kW) each. The WV-2 was later redesignated EC-121K, and numerous specialized versions for the Navy and the Air Force were derived from it.
A dome that rotated
The WV-2E was one of the most unusual. According to the list of Constellation variants, it was an experimental version of the WV-2 modified to carry a rotating radar radome, similar to the one that decades later would make the Boeing E-3 Sentry famous. It's the concept that today identifies almost any airborne early warning aircraft: a large disc mounted on top of the fuselage that rotates with the antenna. The WV-2E was later redesignated EC-121L.
A forerunner in a long-lived family
The WV-2E coexisted with other variants of the same airframe: the WV-2Q, equipped for electronic warfare, later EC-121M; the RC-121D, with fuel tanks on the wingtips, later EC-121D; and the EC-121D itself, the early-warning version of the Big Eye, College Eye, and Disco programs. The Warning Star family served in the United States Navy and Air Force until 1978, more than forty years after work began on the first L-049. That a four-engine piston aircraft born as an airliner, with its characteristic triple tail, should trial the rotating radome gives an idea of just how versatile Lockheed's design turned out to be. That first L-049, the prototype with the civil registration NX 25600, had flown on January 9, 1943, between Burbank and Muroc Field.
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